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Meanwhile, immunofluorescence staining revealed that the number of GRP78 positive cells was significantly increased in the TBI group, and vitamin B12 reversed this trend to a differing degree (Figure 3B)

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The TOX-mediated transcriptional program resulted in severe T-cell exhaustion and upregulated inhibitory immune checkpoint receptors such as PD-1, TIM-3, LAG-3, and TIGIT and reduced the proliferation of cytokine production by CTLs