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The deacetylation of histone lysine residues mediated by HDACs makes chromatin transcriptionally repressive, interfering with gene expression by inhibiting the access of transcription factors (194, 195)

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During glutaminolysis, glutamate is further converted to 2-oxoglutarate by glutamate dehydrogenase, which then enters the tricarboxylic acid (TCA) cycle to form malate and oxaloacetate [10]